Daphnia

__What are daphnia?__

Heres a video on Daphnia, I wasn't able to post it on the website. sorry for any incovinces.

http://videos.howstuffworks.com/hsw/8628-life-in-a-drop-of-water-daphnia-video.htm?sort=company&page=314# [4]



Daphnia are bio indicators that tell if water is healthy or not. When using daphnia for biological tests, water is healthy when there are a good amount of living daphnia. When there are little amount of daphnia, or alot are dead then the water probably is unhealthy or has something wrong with it. [1]

Daphnia are sensitive creatures and can survive only in healthy streams,lakes, and other bodies of water. Because of their sensitivity, they are commonly used as bio indicators. If they don't survive, that almost always means the stream is unhealthy or has some form of pollution in it. [2]

"Animal: Daphnia and/or Water Flea Kingdom: Animalia

Phylum: Crustacea

Class: Branchiopoda

Order: Cladocera

Family Daphniidae

Genus and Species: Daphnia pulex

"The Daphnia is a tiny, pot-bellied creature with a flat back, a rounded head and belly, two black eyes. This tiny creature is common to most healthy ponds, lakes, pools, springs, and streams around the world.

The Daphnia's body is covered by a clear shell, which ends in a point of its tail. The Daphnia's head is not covered the shell. Growing to a length of less than 1/4 inch (four millimeters), the daphnia is one of the tiniest branchiopods. Although the daphnia is very small, it is easy to recognize under a microscope because of its quick, jerky movements. The daphnia moves through the water by the help of its antennae and its few, small, feathery legs.

Like other branchiopods, the daphnia is a foot breather. This activity is what gives this class the name branchiopods, or gill-legs. The Latin word branchio means to breathe, and poda means legs. The water flea breathes through its feet through gill plates which serve as respiratory organs. The daphnia feeds by filtering the water. It beats the water with its small, feathery appendages and pushes tiny particles of organic life into its mouth groove.

The daphnia may reproduce either sexually or asexually. Sexual reproduction involves both the male and the female daphnia. After mating sexually, the female daphina carries her eggs with her in a pouch along the side of her body. The eggs grow and develop there, eventually hatching into miniature daphnia. They are then released from the pouch into the water. Asexual, or independent, reproduction in the daphnia does not include the male daphnia. The female daphina reproduces by a method known as parthenogenesis. Parthenogenesis means the development of an egg without fertilization. It is common for female daphnia to reproduce by parthenogenesis for many generations without the presence of any male daphina.

The daphnia is threatened by a variety of fish and other aquatic, or water-living, animals. In the Great Lakes there are so many daphnia that they have become the main food supply for fish living in those waters.

The daphnia is also often studied by students in middle school and high school life science and biology classes. Since the daphina is common in most ponds and lakes, it is easy to get a daphnia specimen from a one-gallon pond sample. Like many other branchiopods, the daphnia is sometimes sold by humans as fish food.

The life span of the daphnia is not known.

Daphnia are very small and unusual looking creatures. They grow less then 1/4 inch's long, and it's body is covered in a carapace. The daphnia breathe trough their legs, meaning it's gills are on it's feet. They also feed through their feet. Daphnia reproduce sexually, with a male and female, and asexually, just the female. Daphnia are the main food supply for fish in the Great Lakes area. Daphnia are commonly studdied in middle and high school classrooms. They are very easy to get from water samples, and are sold as fish food. How long a daphnia may live is unknown. "[3]

"Geographic Range
Daphnia pulex is the most common species of the water flea, an organism which can be found in almost every permanent, eutrophic (nutrient-rich) water body. A few species are marine, but generally //Daphnia//, including Daphnia pulex, are freshwater organisms.

Habitat
//Daphnia// can be found in almost any permanent body of water, even in rain-filled tire ruts or several meters from the ground, growing in tree moss in a rainforest. They are mainly freshwater and the highest concentrations of //Daphnia// populations are found in the vegetation in most lakes and ponds. They are often the most abundant organism in a body of water. They live as plankton in the open water of lakes, or live either attached to vegetation or near the bottom of the body of water. While very prolific in most freshwater bodies, //Daphnia// are too small and weak to live in a strong current, which they are unable to swim against. They live in a water column and are light enough to stay suspended by using their legs and antennae for movement. They live mainly in the upper portion of this water column near the algae-rich surface of the water, but they will often move up or down the column depending on seasons or predators in a process called diel vertical migration. They are often forced to expend a large amount of energy moving towards a lower depth during the day in order to avoid predators and coming towards the surface to eat at night. Their location is also controlled by seasonal variation in their phytoplankton food supply.

Physical Description
Daphnia pulex is the most common species of the group of organisms known as water fleas. Their common name was given because of their general appearance and jerky swimming motions which resembles that of the land flea. They are, in reality, a type of small crustacean and are generally 0.2-3.0 mm long. Their bodies are not distinctly segmented, but an important feature of their anatomy is the carapace, a folded shell-like structure which covers the animal and opens both ventrally and posteriorly. Studying the anatomy of this organism is made easier by the fact that most of its outer covering is clear, showing most of the internal organs at work, including the heart. The head of the organism contains both a darkly colored compound eye and numerous antennae used for feeling and swimming. Many //Daphnia//, including D. pulex and //D. magna// have a specialized light-sensing organ similiar to a tiny eye called an ocellus. Located posteriorly at the junction of the head are small, hard to see mouthparts. They mainly consist of the mandibles which are in constant motion and used by the organism to crush and grind its food. In a live specimen food particles can be seen passing through the intestine which terminates at the anus located on the postabdomen. The postabdomen is the most posterior part of the body and terminates itself in two hooklike cuticular claws used by the organism to clear debris out of the carapace. The fine teeth located on these claws are often used for species identification. The central portion of the body is the thorax and contains four to six pairs of flattened legs covered in setae. //Daphnia// males are generally smaller than females but have longer antennules and a modified postabdomen. //Daphnia// females posses a brood chamber located between the body wall and dorsal surface of the carapace used to carry their eggs.

Reproduction
Daphnia pulex reproduces both sexually and asexually in a process called parthenogenesis, where male gametes are unnecessary. Parthenogenesis occurs mainly in the summer, so that during summer an entire population of Daphnia pulex will consist almost completely of females. This process begins in the female, which then molt the carapace to increase their size and develope anywhere from two to twenty eggs in their brood chamber. Even without fertilization from a male, these eggs will develope into immature females which are released after the next molting stage. The young that are produced in this way are more precocial or well-developed than in the process of producing altricial fertilized eggs. This stage of reproduction is most used for a rapid increase in //Daphnia// growth but requires more favorable conditions. The sexual stage of //Daphnia// reproduction occurs mainly in the winter during less favorable conditions caused by overcrowding, accumulation of wastes, lower food availability, and lower temperatures. First, some of the eggs that were produced by parthenogenesis hatch into males instead of females. These males then copulate with the females to form fertilized eggs which are then kept in the female's brood chamber. After the female's next molt she releases these eggs which have the ability to overwinter. They can resist freezing and drying while encased in a purselike ephippium that protects the egg as it rests in the sediment at the bottom of the water body until spring. These eggs remain in this stage of arrested developement, lasting up to twenty years, until the conditions become more favorable for hatching. //Daphnia// usually live about ten to thirty days and can live up to one hundred days if their environment is free of predators. An individual will generally have ten to twenty instars, or periods of growth, during their lifetime.

Behavior
Despite the fact that //Daphnia// is a small crustacean, it has received the common name of the water flea because of its close resemblance to that of the terrestial flea. This similarity stems both from their general body structure, which is flattened like that of a flea, and their way of moving through the water by their antennae in a jerky, hopping motion. Many types, however, spend the majority of their time creeping along the bottom of a pond or lake, looking for food particles in the mud. Like all crustaceans, //Daphnia's// shell, or carapace, cannot grow so they are forced to molt as the animal grows larger. The carapace is used for protection and so a new shell is usually grown under the old in order for the organism to be shielded at all times. The old carapace is discarded and the animal rapidly absorbs water into the new shell, producing a stage of very rapid growth called an instar. This is especially prominent in juveniles, which can double their size from one instar to the next. The absorbed water is later gradually replaced by tissue. //Daphnia// are also able to avoid predators in a process called cyclomorphosis in which they change their size and shape in order to be a less suitable foodsource. Their size is often predator specific as they become larger or smaller to avoid consumption by the many types of fish which use //Daphnia// as their primary food. A decrease in size is most common when levels of adult fish population are high, making the //Daphnia// harder for the larger organisms to see; an increase in size is most common when the levels of juvenile fish are high, making the //Daphnia// more difficult for these smaller fish to eat. However, this speciality does come at a reproductive cost to the //Daphnia//.

Food Habits
//Daphnia// are oftened used to clear fish tanks of algae "bloom" because of their diet of bacteria, fine detritus, and very small algae particles. They are filter feeders meaning they do not usually actively seek food; they merely create a constant movement of water using their thoraic legs through their carapace where they are able to filter out any food particles with the setae and direct these towards the mouth. If a mass of food becomes entangled in the mandibles it is cleared by the spines located on the first legs and then kicked out of the carapace by the postabdomen. Not all algae is eaten by //Daphnia//, such as blue-green algae which has too tough of an outer cell wall and filamentous green algae which can be detrimental to the organism's health. While most species of //Daphnia//, including D. pulex, are herbivorous or detritivorous (feeding on phytoplankton), a few are carnivorous and prey on other water fleas.

Economic Importance for Humans: Negative
In general, //Daphnia// are beneficial to an aquatic environment, but they will occasionally limit the population size of other organisms as they compete for food and oxygen. Although they are often used in fish tanks to clear the water of algal bloom, fish are not able to be kept in the same tank with a high number of //Daphnia// because of a limit on the oxygen availability.

Economic Importance for Humans: Positive
Although //Daphnia// are not used by humans as a food source directly, they are involved in many of the foodchains necessary to sustain fish that we consume or use commercially such as sticklebacks, minnows and young Sockeye salmon. They also are a primary food supply for those animals that trout and many other popular fish depend on. Also, almost any freshwater ecosystem is dependent on //Daphnia's// ability to convert phytoplankton and decaying matter into a more usable form. //Daphnia// are also very frequently used by scientists for experimentation. They are small, cheap, and very easy to keep alive in a laboratory environment. Their almost transparent shell makes their internal functions easier to study and they are very susceptible to changes in temperature, food supplies, or dissolved oxygen content in their environment. Aquarium owners often use //Daphnia// both as a food source for their fish and to clear the water of debris.

Conservation Status
//Daphnia// are extremely widespread and common throughout the world. However, they are often used as a food source for aquarium fish and although some of these are raised specifically for this purpose, many are harvested from lakes or ponds. While this practice is unlikely to erradicate all //Daphnia// species, it could damage some rare populations with a limited range.

Other Comments
Genetic variation is often very hard to moniter in //Daphnia// due to their unusual reproductive method of parthenogenesis. Parthenogenic males are difficult to find at certain times, and hybrids will result in a problem as they are often unable to breed or create viable offspring. //Daphnia//, however, still have a very high degree of genetic variation even within a single population or species. They are able to change their size and shape in response to their environment and this ability makes it harder to classify these organisms into specific groups. Often there seems to be more variation within a species than between them." [5]



http://www.mblaquaculture.com/assets/images/content/photo_Daphnia_magna.jpg


 * __References

[1] Mikovich,Ashley.Detective Notebook.19,Nov.,2008 [2] Hellmann,Kaitlin, Detective Notebook. 19, Nov..2008__** [3] [|http://web.ebscohost.com/ehost/detail?vid=3&hid=6&sid=cda017a9-cc9c-4f34-980e-3d622919f22f%40SRCSM2&bdata=JnNpdGU9ZWhvc3QtbGl2ZQ%3d%3d#db=ani&AN=9500200681__**] [4] daphnia video, hellmann, kaitlin, howstuffworks, [5]http://animaldiversity.ummz.umich.edu/site/accounts/information/Daphnia_pulex.html